Residency at University of Florida

• 1982 – 1984, Small Animal Internal Medicine

Past Professional Activities at University of Pennsylvania

• 1984 – 1985, Instructor in Small Animal Medicine
• 1985 – 1990, Assistant Professor of Medicine and Medical Genetics
• 1989 – 2018, Director of the Metabolic Genetics Laboratory - PennGen
• 1989 – present, Secondary Appointment as Privatdozent/Professor at University of Zürich
• 1989 – 2009, Medical Director of Transfusion Medicine Center
• 1990 – 1995, Associate Professor of Medicine and Medical Genetics
• 1990 – present, Secondary Appointment as Professor of Hematology, School of Medicine
• 1995 – 2007, Professor of Medical Genetics and Chief of Medical Genetics
• 1997 – present, Charlotte Newton Sheppard Endowed Professor of Medicine
• 1998 – present, Director of PennGen Laboratories

Honors and Awards

• 1984 – Phi Zeta National Veterinary Honor Society
• 2002 – Scientific Lifetime Achievement Award from the World Small Animal Veterinary Association
• 2002 – Transfusion Medicine Academic Award
• 2004 – John E. McCoy Award from Washington State University for excellence in scientific studies
• 2007 – International Bourgelat Award from British Small Animal Veterinary Association for outstanding contributions to the field of medicine
• 2008 – 2018 WSAVA Hereditary Disease Committee Chair
• 2015 – AVMA Excellence in Feline Research Award for outstanding contributions to feline medicine
• 2018 – Ramsey Distinguished Lecture Award from Iowa State University for clinical research

Current Research/Clinical Interests

His clinical and research expertise and interests are in clinical hematologic disorders, transfusion medicine, hereditary diseases and genetic predispositions in companion animals. They are reflected in >250 original research publications as well as many more reviews and scientific abstracts.  He is a frequently invited speaker at national and international conferences.

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